Decapoda - Brachyura - Xanthidae - Cyclodius
| Cyclodius ungulatus (H. Milne Edwards, 1834) |
World Distribution
Serène (1984 : 252) - Indo-ouest Pacifique. Dans l'océan Indien occidental, Madagascar, Maurice, Réunion, Diego Garcia. Polynésie française. Comprend, Mer Rouge, Afrique orientale, Somalie, Mozambique, Europa, Seychelles, Hawaii.
Environnement : Marine - Substrat/Association : Hard bottom (rock and rubbles)
Vertical range : Shallow-waters (0-100 m) - Min-Max observed: 0-20 m
Mayotte data
Cyclodius ungulatus (H. Milne Edwards, 1834)
Material Examined — Mayotte KUW 2009, st. 12, La Prévoyante Reef, 6-11 m, 4 juv, MNHN-IU-2009-3241, det. July 2011; st. 19, islet Handrema, north, 6-10 m, 1 M 11 x 14 mm (photo), plus 1 sp., MNHN B32444, MNHN-IU-2009-1206; st. 25, islet M'tzamboro, south, 15-20 m, 3 MM 7.95 x 10.95 to 9.55 x 12.90 mm (CW/Lc = 1.36), 1 M 11.1 x 15.1 mm, 1 F 10.1 x 14.1 mm, 3 juv. MNHN-IU-2009-3242, det. July 2011 (see Remarks).
Other records — Phymodius ungulatus - Serène, 1984: 251, fig. 158, 161, pl. XXXV E, Madagascar, Europa, Seychelles; Glorieuses, Island du Lys, intertidal, coll. A. Crosnier, 29 January 1971, 1 M 11.6 x 16.9 mm, 3 FF largest 9.8 x 14.0 mm, MNHN-B6739; Mayotte, intertidal, coll. A. Crosnier, August 1959, 2 MM 7.2 x 10.0 and 8.6 x 12.1 mm, 7 FF largest 10.6 x 15.3 mm, MNHN-B8082. - Poupin et al., 2013c: 11, Glorieuses [Expedition Glorieuses December 2012, st. BIO 39, 1 F 12.9 x 18.9 mm, MNHN-IU-2013-7353].
Remarks — The specimens were determined by using Serène (1984: 247) key with: carapace narrow (CW/Lc<1.5 mm), ambulatory legs with sparse setae, male abdomen with telson and 6th segment as large as long and the male pleopod with long distal setae, directed backward. Cyclodius ungulatus ressembles C. obscurus but is separated by sharper tubercles on the chelae (reduced or absent in C. obscurus).
Geographic and deep ranges — IWP, Red Sea, Mozambique to Hawaii, French Polynesia; intertidal to 15-20 m (KUW fieldwork).
Europa data
Phymodius ungulatus - Dérijard, 1966 : 168, Europa. - Serène, 1984 : 251, fig. 158, 161, pl. XXXV E, Europa, zone intertidale, coll. P. Fourmanoir, 2 MM 7,4 x 10,3 mm et 7,7 x 11,2 mm, MNHN B-6738.
Réunion data
Phymodius ungulatus - Peyrot-Clausade, 1977 : annexe, p. 27 (Réunion). - Ribes, 1978 : 127 (Réunion). - Serène, 1984 : 251 (Réunion d'après Ribes, 1978). - Cyclodius ungulatus - Ng et al., 2008 : 197 (Liste des noms valides). - Mission Poupin, avril 2008 (St. 4, La Saline les Bains, Trou d'Eau, photographies J. Poupin, programme BIOTAS).
New Caledonia data
Cyclodius ungulatus (H. Milne Edwards, 1834) - Juncker & Poupin, 2009, CRISP Report: 66 - Profondeur : 0-6 m / Source : A Milne-Edwards, 1873 - Nouvelles Archives MNHN 8 / Lieu-dit : Lagon Synonyme : Phymodius ungulatus (H. Milne Edwards, 1834).
Central Pacific data (Wallis & Futuna, Polynesia, Clipperton, Easter Island)
Chlorodius ungulatus - Dana, 1852b: 205; 1855, pl. 11, fig. 8a-b (Tahiti). - Chlorodius dehaanii - Heller, 1865: 19 (Tahiti) pro parte fide Forest & Guinot (1961: 110), not C. Dehaani, synonym of Phymodius granulatus (Targioni Tozzetti, 1877) in Serène (1984: 250). - Cyclodius gracilis - Nobili, 1907: 397 (Fakahina, "Rikitea" = Mangareva). - Phymodius ungulatus - Rathbun, 1907: 46, pl. 3-4 (Bora Bora, Fakarava, Makemo, Rangiroa, Tahiti) pro parte fide Forest & Guinot (1961: 110; cf. Remark). - Nobili, 1907: 393 ("Rikitea" = Mangareva). - Boone, 1934: 140, pl. 73 (Raiatea, Tahiti) pro parte fide Forest & Guinot (1961: 110). - Holthuis, 1953: 25 (Raroia). - Forest & Guinot, 1961: 110, fig. 86a-b, pl. 11, fig. 1-4, pl. 12, fig. 1-4, pl. 13, fig. 1-3, pl. 14, fig. 1-3 (Fakahina, Hikueru, Mangareva, Tahiti); 1962: 66 (Biogeography "Tahiti-Tuamotu"). - Guinot, 1964: 74, fig. 38 (Tahiti); 1985: 451 (List). - Peyrot-Clausade, 1977a, annex of the species: 27; 1977b: 213 (Moorea); 1989: 113 (Tikehau). - Monteforte, 1984: 171, annex 1, tab. a (Mataiva, Moorea, Tahiti, Takapoto); 1987: 9 (Moorea). - Serène, 1984: 251, fig. 158, 161, pl. 35e (Tahiti; Syn.). - Delesalle, 1985: 288 (Mataiva). - Odinetz-Collart & Richer de Forges, 1985: 201 (Moorea and/or Tahiti, Takapoto). - Salvat & Richard, 1985: 356 (Takapoto). - Poupin, 1996a: 55 (Checklist) - New material - Coll. C. Hily, det. J. Poupin ("Arue" = Tahiti); coll. Frv Marara, det. J. Poupin (Fangataufa). - Synonyms - Cyclodius gracilis Dana, 1852. - Remark: Forest & Guinot (1961: 111) indicate that Rathbun's material is only partially assignable to this species, but without mentioning the localities referring to the real Phymodius ungulatus. Thus, for this species, some of the islands mentioned by Rathbun are doubtful. - Unpublished specimens, coll. & det. G. Paulay, Tikehau, June 10, 2001, Tuherahera Motu, Lagoon fringing reef and sand flat in front to Aito Motel (FLMNH-UF 1572, 1461). Cyclodius ungulatus - Ng et al., 2008 : 197 (List of valid names). - Moorea Biocode project. - Mission CRIOBE 2014 (collection, Moorea, 1982, coll. & det. Monteforte).
References : PDF list, 214 pp (1.4 Mo)This species in Worms Database